By Robbie Whelan 

BELLEVUE, Wash.--The nation's high-end malls are posting record sales, thanks in part to the growing presence of technology retailers that sell pricey goods.

At Bellevue Square, a shopping center in this affluent Seattle suburb, electric car maker Tesla Motors Inc. produced sales of $5,500 per square foot last January, more than five times the mall's average and at a pace the mall's owners say is likely a U.S. record for any retailer.

At a time when some big department stores are struggling and Internet shopping is on the rise, the mall industry is doing surprisingly well. Sales have risen each year since the recession, from $383 per square foot in 2009 to an estimated $478 in 2014, says the International Council of Shopping Centers.

Malls owned by real-estate investment trusts, which tend to have more luxury tenants and often are located in large population centers, generated a record $550 in per-square-foot sales in the third quarter of 2014, shattering the previous high of $450 a square foot reached in 2007, according to REIT research firm Green Street Advisors.

No one tracks sales per square foot across all retail stores. But landlords say a big driver of malls' recent resurgence has been a new generation of technology-focused tenants, such as Tesla, Apple Inc. and Microsoft Inc., that are ringing up strong sales and supplanting "anchor" department stores that malls used to rely on to bring in customers. Tesla and Apple declined to disclose figures for their stores. Microsoft couldn't be reached for comment.

"They do very high volumes," said Peter Lowy, co-chief executive of Westfield Corp., of the high-tech tenants. Westfield has 16 Apple stores, six Tesla locations and six Microsoft stores in its 38 U.S. malls. "Your company's sales per square foot are really boosted by those tenants."

General Growth Properties Inc., the nation's second-largest mall operator, said Apple's rollout of the iPhone 6 was a main driver of its 6.7% increase in September monthly sales. Without Apple, sales would have risen only 4%, Chief Executive Sandeep Mathrani told analysts in October.

Some traditional mall anchors like J.C. Penney Co. and Sears Holdings Inc. are in a prolonged slump. J.C. Penney suffered a per-square-foot sales decline of 30% from 2010 to 2013, the most recent year for which data are available, according to investment bank Imperial Capital. Sears doesn't report average sales per square foot, but sales at stores open at least a year have fallen by a cumulative 12.1% since 2010, according to investment bank Evercore ISI.

Yet for malls overall, the trend is moving the other way: Sales per square foot among mall REITs are up 36% since 2010, according to Green Street. That is a sign that mall owners are pruning their portfolios of less-productive properties while filling vacant spaces with more-productive retailers, including high-tech tenants.

The new tenants tend to be far more profitable for malls than anchor stores because, under typical lease agreements, they are required to share a percentage of their sales with their landlords.

Anchor tenants, by contrast, typically own their spaces outright, though they often contribute fees to help maintain common spaces. They serve mall owners mainly by attracting shoppers to the smaller retailers between the anchors, known as "inline" stores.

Investors have benefited from malls' resurgence. In all, REITs that own regional malls produced a total return of 35% in 2014, including dividends, according to the National Association of Real Estate Investment Trusts, second only to apartments and health-care properties among the big REIT categories.

High-tech stores have helped malls fend off the Internet challenge as well.

The share of retail sales completed online has risen steadily, from 4% in the third quarter of 2009 to 6.6% in the third quarter of 2014, according to the most recent data available from the National Retail Federation.

One reason malls have held steady: Merchants such as Tesla, Apple and the like "show how the malls have stayed very relevant," says Alexander Goldfarb, a REIT analyst with Sandler O'Neill + Partners.

High-tech tenants are a recent phenomenon. Over the past decade, the number of Apple stores in North American malls has grown to 219 from just a handful. Microsoft has opened 70 store locations in malls since 2009 and Tesla 25 since 2008, according to Green Street.

In Bellevue, the affluent suburb east of Seattle filled with large houses and a central business district with a host of shiny new office buildings under construction, Bellevue Square is the city's most prominent shopping center. The town's median household income was $88,073 in 2012, about 73% higher than the national median.

Tesla's $5,500 in sales per square foot last January shows how lucrative high-tech tenants can be.

Yet it wasn't a fluke, says Kemper Freeman, chairman of Kemper Development Co., the family-run company that has owned Bellevue Square since it was built at the end of World War II. The store also was the highest-producing tenant in the mall for the rest of the year on a sales-per-square-foot basis.

Overall, including Tesla, the mall produced $1,071 in sales per square foot for all of 2013, up from $842 in 2012, according to the company. In its best-performing month in 2014, Bellevue Square's Apple store produced about $1,500 per square foot, Mr. Freeman said. Tesla opened its store there in November 2011, a year after Microsoft opened its store.

Nationwide, the highest-quality U.S. malls produced $666 in average sales per square foot in 2013, according to Green Street.

Tesla isn't the only high-tech attraction at the mall. Last January, sales at the mall's 12,000-square foot Apple Store matched the same dollar volume as one of its largest anchors with more than 250,000 square feet of space. And three weeks ago, a store by specialty photography brand Leica opened, selling digital cameras for up to $22,000 apiece.

"In the old days, a few major anchors paid the rent," Mr. Freeman said. "The inline stores are, at this point, subsidizing the main stores."

Karen Busto, an executive at a company in Mexico City that distributes automotive lubricants, visited the Tesla store with a colleague during a shopping break on a business trip. She said the store's mall location made perfect sense to her because it would give the cars more exposure and allow curious customers to stop in and learn more.

"Enthusiasts who love this kind of thing, for them it's fantastic to have this type of store here," she said. "It's more personal than going to a dealership."

Write to Robbie Whelan at robbie.whelan@wsj.com

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