LONDON—Former bank trader Tom Hayes accused top Citigroup Inc. executives of being involved in efforts to manipulate global benchmark rates and then engineering a coverup, a London court heard Monday.

Brian McCappin, the chief executive of Citigroup's global markets business in Japan, "would talk to the cash desk in Tokyo and ask them to move Tibors [Tokyo interbank offered rate],"according to an email sent by Mr. Hayes to a contact at Barclays PLC on Sept. 7, 2010, the day after Mr. Hayes was dismissed from the U.S. bank.

A Citigroup spokesman declined to comment on the trial or the allegations against Mr. McCappin.

Another senior colleague at Citigroup was copied on an email that contributed to Mr. Hayes's firing on allegations of attempted manipulation of the Yen London interbank offered rate, or Yen Libor, and the Tokyo interbank offered rate, or Tibor, Mr. Hayes told the Barclays contact in the email message.

"My superior was cc'd on the email to my junior when I mention talking to them about why they moved the fix," Mr. Hayes said, expressing dismay about losing his job after an internal investigation, "...am still in shock."

In that December 2009 email to the junior colleague, seen by the jury earlier Monday, Mr. Hayes had asked the colleague to find out from rate submitters in London why their rate had changed recently and to get information about future moves so that "if we know ahead of time we can position and scalp the market."

Mr. Hayes is on trial for eight counts of fraud related to alleged manipulation of Libor and has said he isn't guilty. Mr. McCappin hasn't been accused of any wrongdoing by authorities.

The court also heard how Mr. Hayes in a June 2013 interview with British prosecutors said that he was instructed by Mr. McCappin to say that he had only asked a junior colleague to share information with rate submitters in London rather than request that they adjust their submissions.

During the internal investigation, Mr. Hayes wasn't suspended and was discouraged from seeking legal advice, Mr. Hayes told the prosecutors. While Mr. Hayes was dismissed, a senior colleague was allowed to resign, and "Brian's involvement was covered up," Mr. Hayes said in the interview.

Write to Margot Patrick at margot.patrick@wsj.com

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