By Chiara Albanese 

Bank of New York Mellon Corp. has fired a London-based currency trader after the bank found that his electronic communications breached its compliance rules, according to a person familiar with the matter.

The dismissal is the first of its kind from this bank, or from any chiefly custody-focused bank, since the global regulatory investigation into currencies-trading practices started early last year, a sign that behavior is under scrutiny beyond the group of top dealing banks that agreed to pay $4.3 billion in fines on both sides of the Atlantic earlier this month.

Jon Smailes, senior trader on the London spot desk of the bank, was ousted in recent months following a period of leave, based on the findings of an internal review initiated by BNY Mellon around the time that regulators started digging into the market last year, according to the person. His departure hasn't been previously reported. Mr. Smailes, who had been at the bank for more than 10 years, couldn't be reached for comment. The nature of his communications and what rules he is said to have broken are unclear.

The bank informed U.K. regulator the Financial Conduct Authority of its findings and of Mr. Smailes's dismissal, the person familiar with the matter said. The bank isn't under investigation in the U.K.

New York-headquartered BNY Mellon is a relatively small player in the $5.3 trillion-a-day currencies market, and the bulk of its foreign-exchange business stems from its activities in custody, the safekeeping of assets for clients such as pension funds. The bank has about $28.3 trillion of assets under custody or administration, according to its website.

The bank wasn't one of those fined earlier this month for their failure properly to supervise traders. Citigroup Inc., J.P. Morgan Chase & Co., Royal Bank of Scotland Group PLC, HSBC Holdings PLC, Bank of America Corp. and UBS AG reached settlements with a combination of regulators in the U.K., U.S. and Switzerland.

Regulators said those banks failed to stop employees from improperly sharing confidential information with rival banks or from improperly attempting to boost currencies-trading profits at their customers' expense. The banks didn't dispute the regulators' findings.

The Nov. 12 settlement hasn't proven to be the end of the matter. Tuesday, The Wall Street Journal reported that the U.S. Justice Department is investigating allegations that an employee of HSBC leaked confidential client information to a major hedge fund.

Last week, Goldman Sachs Group Inc., another firm that wasn't part of regulatory settlements earlier this month, also fired a currencies trader who had previously held a role at HSBC.

Following the settlement, RBS Chief Executive Ross McEwan said his bank will continue its internal investigation of its foreign-exchange business. At the time, he said RBS is examining disciplinary and accountability processes for more than 50 traders and employees. "Those who have been found lacking in conduct or accountability terms will be dealt with appropriately, including through clawback, award forfeiture, or through formal disciplinary procedures," he said.

All told, about a dozen banks have so far suspended or fired more than 30 traders and other employees in connection with the foreign-exchange probes.

Write to Chiara Albanese at chiara.albanese@wsj.com

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