By Ryan Knutson 

RALEIGH N.C.--Of all the eyes on Google Inc.'s new wireless service last week, none were watching more intently than those at the North Carolina headquarters of Republic Wireless--a startup that is about to face a big new rival.

Google's new service, called Project Fi, takes the potentially disruptive approach of routing users over free Wi-Fi airwaves where possible and using expensive cellular service only as a fallback. That's exactly what Republic has done as it has built up a service with a few hundred thousand subscribers over the past four years.

It isn't clear what the entrance of a big consumer brand like Google means for Republic. Optimists say it validates the Wi-Fi-first approach and may help Republic persuade subscribers to switch from mainstream carriers while expanding the number of phones that will work on the service. But there's also a risk that Google could end up disrupting the 225-person company that moved more quickly to challenge the telecom industry's predominant pricing.

Google and Republic have a relationship that dates back years, and at one point employees at Google's former Motorola unit brainstormed about acquiring Republic. Now, though, Google has jumped into Republic's space. In a sign of the fresh competition, Republic rolled out a plan last week to charge subscribers only for the data they use, and Google's service was unveiled with an identical concept just two days later.

"It's like having a famous person move next door," said Republic's co-founder, David Morken. "You want to give them a warm welcome and hope they know how to be a good neighbor."

Google declined to comment.

A few other companies also offer a Wi-Fi first approach, including the startups Scratch Wireless and FreedomPop. Cable companies have also been exploring the Wi-Fi-first idea, and Cablevision has launched a Wi-Fi-only service.

Republic Wireless is a unit of Bandwidth.com, a company that Mr. Morken founded in 1999 to resell Internet access to small businesses. In 2001, Republic relocated to North Carolina and built a network to route calls over the Internet that is now used by Skype and Vonage. In 2006, it began working with Google to help its Gmail voice service, now known as Google Voice, make calls to regular phones.

The wireless venture came about after Mr. Morken promised his oldest child an iPhone if he got straight A's. He then extended the promise to the rest of his children. By the time the fourth oldest won a phone, Mr. Morken's AT&T bill had topped $1,000 a month.

Mr. Morken wondered if he could use Bandwidth.com's Internet phone technology to offer cheaper wireless service by leaning on the Wi-Fi hotspots that many people already have at home, at work and at sites like airports. Republic began selling its service in 2011 with just 6,000 test phones. In the first 24 hours, about 100,000 people tried to sign up.

A key challenge was to figure out how to keep calls from dropping when users moved out of a hotspot. Republic found an answer in 2013: When the Wi-Fi signal gets weak, the device begins a second call over the cellular network and then merges the calls. The company has more than a dozen patents related to the capability, which is known as seamless handover.

Republic sells service with cellular-voice-and-text fallback starting at $10 a month. The base plan for Google's new offering costs $30. Republic has around 300,000 subscribers.

The handover feature on Google's Project Fi works similarly. Wi-Fi-calling features available from Scratch Wireless, T-Mobile and Sprint don't yet have the same seamless-handover function.

Jonathan Chaplin, a telecom analyst at New Street Research, said Google's entry will help Republic because it validates the Wi-Fi-first concept. Google's size may also encourage more phone makers to support the technology on their devices, thus broadening the number of phones Republic can sell.

Right now, customers can use Republic on phones made by Motorola, which has worked closely with Republic to make the service work. After Google bought the phone maker in 2012, Motorola employees were given a challenge: Figure out how to sell one billion phones.

Motorola employees discussed internally whether acquiring Republic might help achieve that goal, according to a former employee involved in the discussions. The thinking was that if Motorola could sell phones with Republic's inexpensive wireless service, they would be more attractive.

Google declined to comment on the discussions. The idea was dropped after Google sold Motorola to Lenovo Group Ltd. in 2014.

Mr. Morken is an avid cyclist who has completed two 100 mile ultramarathons, but he also believes in time off. A few days a week, he circles the office and encourages employees to go home. When workers go on vacation, they are barred from communicating with anyone about work.

In 2012, Mr. Morken was about to leave on a two-week trip to Wyoming with his family when he learned that Republic's warehouse owner in Texas was going bankrupt with Republic's phones locked up inside. He got the ball rolling on a solution, then left for his vacation anyway. Mr. Morken said that going away turns younger staffers into leaders by forcing them to make tough decisions.

"I had no idea whether when I got back whether we'd still have a Republic Wireless," he said.

Alistair Barr contributed to this article.

Write to Ryan Knutson at ryan.knutson@wsj.com

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